I'm really curious how the gains are calculated now and into the future.
Suppose for example I owned 1,000 shares of the index mutual fund VTSMX at $60/share, with a cost basis per share of $50/share giving unrealized gains of $10/share ($10,000). I now convert it to the ETF equivalent, VTI.
Vanguard do this with no US tax implications and now I own 500 shares at $120/share, with cost basis of $100/share, (these are the approximate current prices) with unrealized gains of $20/share ($10,000).
Will HMRC consider that I have actually sold the VTSMX shares and owe tax on a capital gain of $10,000? If so then I will need to keep my own records and know that my cost basis on date of conversion is $120/share. Ongoing the cost basis of my VTI shares is going to be different for the IRS and HMRC.
ETA
If this is the case then it would be better to actually sell the VTSMX shares and then buy the VTI shares. That way the HMRC tax can used as a credit against the US taxes and ongoing the accounting will be much easier.
As VTSMX is a non-reporting fund I think you will have to use every transaction and the price at that point to work out the gains on the fund. If you have been reinvesting dividends or making regular small purchases this gets very tedious, even if you have kept detailed records. But let's say you have that information and work out the capital gain. It looks like Guya is saying that the conversion from VTSMX to VTI will trigger a UK tax event even though the underlying investments are exactly the same. If that's the case you will pay tax on the gains at your marginal income tax rate.
If you did the conversion before being subject to UK taxation and then sold VTI while a UK resident you would have a capital gain and Vanguard would send you a 1099 to show that. I tell Vanguard to use the average cost basis when calculating gains because it's simple and if you do that you can use those numbers on your HMRC tax bill. As VTI is a reporting fund you can use the capital gains tax free allowance and pay at capital tax gains tax rates.