1) Applying for UK citizenship alone is not enough to establish domicile. It depends in part on why you are applying (which you must put on your application). If you want to be sure you phrase everything in such a way as to ensure you don't lose your domicile by obtaining citizenship, then use the services of a lawyer or immigration specialist whose job it is to know what to say and how to say it, and who can be held accountable for screwing up.
2) You gave away way, way, way too much information on your post. Give any further info offline in a telephone or in-person conversation with your advisor.
3) Trusts may in fact have UK tax as well. (Don't reply here!) If the trustees are resident in Britain, it turns it into a British trust. Hopefully, your trustees are NOT living in the UK, and if they are, HOPEFULLY you can retroactively change them to non-UK-resident persons. Also, any remittances from the trust into the UK are taxed in the UK (less any UK tax paid).
4) Pensions and investment income are taxed in the UK if you remitted them to the UK. If you kept all this money offshore, then you did the right thing taxing them only in the US.
5) You can restructure your investments in such a way that there's no UK tax. This is because after your restructuring, the monies you are remitting to the UK are considered capital instead of investments, pensions, or trust proceeds. That requires some serious work between your dual US/UK tax consultant and your US financial planner.
6) As you can see, you've opened a mega can of worms by including way too much information that was irrelevant to your actual question. I wouldn't ask these kinds of questions here, and wherever you do ask them, don't do it in writing and use the phrase "hypothetically speaking" a lot. A lot. Exclusively, even. Why? So that you can figure out what you need to do without committing yourself to an immediate plan of action. Not paying taxes is a crime in the UK, so you have to resolve it quickly once you've brought the problem to the attention of a professional. Thus, hypothetically speaking gives you time to take care of things at your own speed. You may be able to make some changes retroactively, and thus you may be able to significantly reduce any UK tax exposure. Get serious, real, offline advice.